WHITE AND GREENISH FLOWERS
“The transition from wind-fertilization to insect-fertilization and the first traces of adaptation to insects, could only be due to the influence of quite short-lipped insects with feebly developed color sense. The most primitive flowers are therefore for the most part simple, widely open, regular, devoid of nectar or with their nectar unconcealed and easily accessible, and greenish, white, or yellow in color…. Lepidoptera, by the thinness, sometimes by the length, of their tongues, were able to produce special modifications. Through their agency were developed flowers with long and narrow tubes, whose colors and time of opening were in relation to the tastes and habits of their visitors. ” - Hermann Muller.
“Of all colors, white is the prevailing one; and of white flowers a considerably larger proportion smell sweetly than of any other color, namely, 14. 6 per cent; of red only 8. 2 per cent are odoriferous. The fact of a large proportion of white flowers smelling sweetly may depend in part on those which are fertilized by moths requiring the double aid of conspicuousness in the dusk and of odor. So great is the economy of Nature, that most flowers which are fertilized by crepuscular or nocturnal insects emit their odor chiefly or exclusively in the evening. ” - Charles Darwin.
WATER-PLANTAIN
(Alisma Plantago-aquatica) Water-plantain family
Flowers - Very small and numerous, white, or pale pink, whorled in bracted clusters forming a large, loose panicle 6 to 15 in. long on a usually solitary scape 1/2 to 3 ft. high. Calyx of 3 sepals corolla of 3 deciduous petals; 6 or more stamens; many carpels in a ring on a small flat receptacle. Leaves: Erect or floating, oblong or ovate, with several ribs, or lance-shaped or grass-like, petioled, all from root. Perferred Habitat - Shallow water, mud, marshes. Flowering Season - June-September. Distribution - North America, Europe, Asia.
Unlike its far more showy, decorative cousin the arrow-head, this wee-blossomed plant, whose misty white panicles rise with compensating generosity the world around, bears only perfect, regular flowers. Twelve infinitesimal drops of nectar, secreted in a fleshy ring around the center, are eagerly sought by flies. As the anthers point obliquely outward and away from the stigmas, an incoming fly, bearing pollen on his under side, usually alights in the center, and leaves some of the vitalizing dust just where it is most needed. But a “fly starting from a petal, ” says Muller, “usually applies its tongue to the nectar-drops one by one, and after each it strokes an anther with its labellae; in so doing it may bring various parts of its body in contact with the anthers. As a rule, however, the parts which come in contact with the anthers are not those which come in contact with the stigmas in the same flower. ” Any plant that lives in shallow water, which may dry up as summer advances, is under special necessity to produce an extra quantity of cross-fertilized seed to guard against extinction during drought. For the same reason it bears several kinds of leaves adapted to its environment: broad ones that spread their surfaces to the sunshine, and long grass-like ones to glide through currents of water that would tear those of any other shape. What diversity of leaf-form and structure we meet daily, and yet how very little does the wisest man of science understand of the reasons underlying such marvellous adaptability!
BROAD-LEAVED ARROWHEAD (Sagittaria latifolia; S. variabilis of Gray) Water-plantain family
Flowers - White, 1 to 1 1/2 in. wide, in 3-bracted whorls of 3, borne near the summit of a leafless scape 4 in. to 4 ft. tall. Calyx of 3 sepals corolla of 3 rounded, spreading petals. Stamens and pistils numerous, the former yellow in upper flowers usually absent or imperfect in lower pistillate flowers. Leaves: Exceedingly variable; those under water usually long and grasslike; upper ones sharply arrow-shaped or blunt and broad, spongy or leathery, on long petioles. Preferred Habitat - Shallow water and mud. Flowering Season - July-September. Distribution - From Mexico northward throughout our area to the circumpolar regions.
Wading into shallow water or standing on some muddy shore, like a heron, this striking plant, so often found in that bird's haunts, is quite as decorative in a picture, and, happily, far more approachable in life. Indeed, one of the comforts of botany as compared with bird study is that we may get close enough to the flowers to observe their last detail, whereas the bird we have followed laboriously over hill and dale, through briers and swamps, darts away beyond the range of field-glasses with tantalizing swiftness.
While no single plant is yet thoroughly known to scientists, in spite of the years of study devoted by specialists to separate groups, no plant remains wholly meaningless. When Keppler discovered the majestic order of movement of the heavenly bodies, he exclaimed, “Oh God, I think Thy thoughts after Thee! ” - the expression of a discipleship every reverent soul must be conscious of in penetrating, be it ever so little a way, into the inner meaning of the humblest wayside weed.
Fragile, delicate, pure white, golden-centered flowers of the arrowhead, usually clustered about the top of the scape, naturally are the first to attract the attention whether of man or insect. Below these, dull green, unattractive collections of pistils, which by courtesy only may be called flowers, also form little groups of three. Like the Quakers at meeting, the male and female arrowhead flowers are separated, often on distinct plants. Of course the insect visitors - bees and flies chiefly - alight on the showy staminate blossoms first, and transfer pollen from them to the dull pistillate ones later, as it was intended they should, to prevent self-fertilization. How endless are the devices of the flowers to guard against this evil and to compel insects to cross-pollinate them! The most minute detail of the mechanism involved, which the microscope reveals, only increases our interest and wonder.
Any plant which elects to grow in shallow water must be amphibious; it must be able to breathe beneath the surface as the fish do, and also be adapted to thrive without those parts that correspond to gills; for ponds and streams have an unpleasant way of drying up in summer, leaving it stranded on the shore. This accounts in part for the variable leaves on the arrowhead, those underneath the water being long and ribbon-like, to bring the greatest possible area into contact with the air with which the water is charged. Broad leaves would be torn to shreds by the current through which grass-like blades glide harmlessly; but when this plant grows on shore, having no longer use for its lower ribbons, it loses them, and expands only broad arrow-shaped surfaces to the sunny air, leaves to be supplied with carbonic acid to assimilate, and sunshine to turn off the oxygen and store up the carbon into their system.
WATER ARUM; MARSH CALLA
(Calla palustris) Arum family
Flowers - Minute, greenish yellow, clustered on a cylinder-like, fleshy spadix about 1 in. long, partly enfolded by a large, white, oval, pointed, erect spathe, the whole resembling a small calla lily open in front. The solitary “flower” on a scape as long as the petioles of leaves, and, like them, sheathed at base. Leaves: Thick, somewhat heart-shaped, their spreading or erect petioles 4 to 8 in. long. Fruit: Red berries clustered in a head. Preferred Habitat - Cool Northern bogs; in or beside sluggish water. Flowering Season - May-June. Distribution - Nova Scotia southward to Virginia, westward to Minnesota and Iowa.
At a glance one knows this beautiful denizen of Northern bogs and ditches to be a poor relation of the stately Ethiopian calla lily of our greenhouses. Where the arum grows in rich, cool retreats, it is apt to be abundant, its slender rootstocks running hither and thither through the yielding soil with thrifty rapidity until the place is carpeted with its handsome dark leaves, from which the pure white “flowers” arise; and yet many flower lovers well up in field practice know it not. Thoreau, for example, was no longer young when he first saw, or, rather, noticed it. “Having found this in one place, ” he wrote, “I now find it in another. Many an object is not seen, though it falls within the range of our visual ray, because it does not come within the range of our intellectual ray. So, in the largest sense, we find only the world we look for. ”
Now, the true flowers of the arum and all its spadix-bearing kin are so minute that one scarcely notices them where they are clustered on the club-shaped column in the center of the apparent “flower. ” The beautiful white banner of the marsh calla, or the green and maroon striped pulpit from which Jack preaches, is no more the flower proper than the papery sheath below the daffodil is the daffodil. In the arum the white advertisement flaunted before flying insects is not even essential to the florets' existence, except as it helps them attract their pollen-carrying friends. Almost all waterside plants, it will be noticed, depend chiefly upon flies and midges, and these lack aesthetic taste. “Such plants have usually acquired small and inconspicuous separate flowers, ” says Grant Allen; “and then, to make up for their loss in attractiveness, like cheap sweetmeats, they have very largely increased their numbers. Or, to put the matter more simply and physically, in waterside situations those plants succeed best which have a relatively large number of individually small and unnoticeable flowers massed together into large and closely serried bundles. Hence, in such situations, there is a tendency for petals to be suppressed, and for blossoms to grow minute; because the large and bright flowers seldom succeed in attracting big land insects like bees or butterflies, while the small and thick-set ones usually do succeed in attracting a great many little flitting midges. ” Flies, which are guided far more by their sense of smell than by sight, resort to the petalless, insignificant florets of the ill-scented marsh calla in numbers; and as the uppermost clusters are staminate only, while the lower florets contain stamens and pistil, it follows they must often effect cross-pollination as they crawl over the spadix. But here is no trap to catch the tiny benefactors such as is set by wicked Jack-in-the-pulpit, or the skunk-cabbage, or another cousin, a still more terrible executioner, the cuckoo-pint (Arum maculatum) of Europe.
Few coroner's inquests are held over the dead bodies of our feathered friends; and it is not known whether the innocent-looking marsh calla really poisons the birds on which it depends to carry its bright seeds afar or not. The cuckoo-pint, as is well known, destroys the winged messenger bearing its offspring to plant fresh colonies in a distant bog, because the decayed body of the bird acts as the best possible fertilizer into which the seedling may strike its roots. Most of our noxious weeds, like our vermin, have come to us from Europe; but Heaven deliver us from this cannibalistic pest!
The very common GREEN ARROW-ARUM (Peltandra Virginica), found in shallow water, ditches, swamps, and the muddy shores of ponds throughout the eastern half of the United States, attracts us more by its stately growth and the beauty of its bright, lustrous green arrow-shaped leaves (which have been found thirty inches long), than by the insignificant florets clustered on the spadix within a long pointed green sheath that closely enfolds it. Pistillate florets cover it for only about one-fourth its length. To them flies carry pollen from the staminate florets covering the rest of the spadix. After the club is set with green berries - green, for this plant has no need to attract birds with bright red ones - the flower stalk curves, bends downward, and the pointed leathery sheath acting as an auger, it bores a hole into the soft mud in which the seeds germinate with the help of their surrounding jelly as a fertilizer.
AMERICAN WHITE HELLEBORE; INDIAN POKE; ITCH-WEED
(Veratrum viride) Bunch-flower family
Flowers - Dingy, pale yellowish or whitish green, growing greener with age, 1 in. or less across, very numerous, in stiff-branching, spike-like, dense-flowered panicles. Perianth of 6 oblong segments; 6 short curved stamens; 3 styles. Stem: Stout, leafy, 2 to 8 ft. tall. Leaves: Plaited, lower ones broadly oval, pointed, 6 to 12 in. long; parallel ribbed, sheathing the stem where they clasp it; upper leaves gradually narrowing; those among flowers small. Preferred Habitat - Swamps, wet woods, low meadows. Flowering Season - May-July. Distribution - British Possessions from ocean to ocean; southward in the United States to Georgia, Tennessee, and Minnesota.
"Borage and hellebore fill two scenes -
Sovereign plants to purge the veins
Of melancholy, and cheer the heart
Of those black fumes which make it smart. "
Such are the antidotes for madness prescribed by Burton in his “Anatomie of Melancholy. ” But like most medicines, so the homeopaths have taught us, the plant that heals may also poison; and the coarse, thick rootstock of this hellebore sometimes does deadly work. The shining plaited leaves, put forth so early in the spring they are especially tempting to grazing cattle on that account, are too well known by most animals, however, to be touched by them - precisely the end desired, of course, by the hellebore, nightshade, aconite, cyclamen, Jamestown weed, and a host of others that resort, for protection, to the low trick of mixing poisonous chemicals with their cellular juices. Pliny told how the horses, oxen, and swine of his day were killed by eating the foliage of the black hellebore. Flies, which visit the dirty, yellowish-green flowers in abundance, must cross-fertilize them, as the anthers mature before the stigmas are ready to receive pollen. Apparently the visitors suffer no ill effects from the nectar. We nave just seen how the green arrow-arum bores a hole in the mud and plants its own seeds in autumn. The hellebore uses its auger in the spring, when we find the stout, shining, solid tool above ground with the early skunk-cabbage.
STAR OF BETHLEHEM; TEN O'CLOCK
(Ornithogalum umbellatum) Lily family
Flowers - Opening in the sunshine, white within, greenish on the outside, veined, borne on slender pedicels in an erect, loose cluster. Perianth of 6 narrowly oblong divisions, 1/2 in. long or over, or about twice as long as the flattened stamens; style short, 3-sided. Scape: Slender, 4 to 12 in. high, with narrow, blade-like bracts above. Leaves: Narrow, grass-like with white midvein, fleshy, all from coated, egg-shaped bulb. Preferred Habitat - Moist, grassy meadows, old lawns. Flowering Season - May-June. Distribution - Escaped from gardens from Massachusetts to Virginia.
The finding of these exquisite little flowers, growing wild among the lush grass of a meadow not far from some old homestead where their ancestors, with crocuses and grape hyacinths, once brightened the lawn in early spring, makes one long to start a Parkinson Society instantly. Some school children not far from New York, receiving their inspiration from Mrs. Ewing's little book, “Mary's Meadow, ” have spread the gospel of beauty, like the true missionaries they are, by systematically planting in lanes and fields sweet violets, golden coreopsis, hardy poppies, blue corn-flowers, Japanese roses, orange day-lilies, larkspurs, and many other charming garden flowers that need only the slightest encouragement to run wild. Immense quantities of seed, that go to loss in every garden, might so easily be sprinkled at large on our walks. Nearly all the beautiful hardy perennials cultivated here grow in Nature's garden in Europe or Asia, and will do so in America if they are but given the chance. The Star of Bethlehem is a case in point. Several members of the large group of charming spring flowers to which it belongs grow in such abundance in the Old World that for centuries the bulbs have furnished food to the omnivorous Italian and Asiatic peasants. If we cannot spare offsets from the garden, and will wait a few years for seeds to bear, the rich, light loam of our grassy meadows, too, will be streaked with a Milky Way of floral stars, as they are in Italy.
The Greek generic name of the Star of Bethlehem, meaning “bird's milk” (a popular folk expression in Europe for some marvellous thing) was applied by Linnaeus because of the flower's likeness to the wonderful star in the East which guided the Wise Men to the manger where Jesus lay.
STAR-GRASS; COLIC-ROOT
(Aletris farinosa) Lily family
Flowers - Small, oblong-tubular, pure white or yellowish, about 1/4 in. long, set obliquely in a long, wand-like, spiked raceme, at the end of a slender scape 2 to 3 ft. tall. Perianth somewhat bell-shaped, 6-pointed, rough or mealy outside; 6 stamens, inserted below each point; style 3-cleft at tip. (A Southern form or distinct species (? ) has yellower, fragrant flowers. ) Leaves: >From the base, lance-shaped, 2 to 6 in. long, thin, pale yellowish green, in a spreading cluster. Preferred Habitat - Dry soil; roadsides; open, grassy, sandy woods. Flowering Season - May-July. Distribution - From Ontario and the Mississippi eastward to the Atlantic.
Herb gatherers have searched far and wide for this plant's bitter, fibrous root, because of its supposed medicinal virtues. What decoctions have not men swallowed from babyhood to old age to get relief from griping colic! In partial shade, colonies of the tufted yellow-green leaves send up from the center gradually lengthening spikes of bloom that may finally attain over a foot in length. The plant is not unknown in borders of men's gardens. The Greek word (aletron = meal) from which its generic title is derived, refers to the rough, granular surface of the little oblong white flower.
WILD SPIKENARD; FALSE SOLOMON'S SEAL; SOLOMON'S ZIG-ZAG
(Vagnera racemosa; Smilacina racemosa of Gray)
Lily-of-the-Valley family
Flowers - White or greenish, small, slightly fragrant, in a densely flowered terminal raceme. Perianth of 6 separate, spreading segments; 6 stamens; 1 pistil. Stem: Simple, somewhat angled, 1 to 3 ft. high, scaly below, leafy, and sometimes finely hairy above. Leaves: Alternate and seated along stem, oblong, lance-shaped, 3 to 6 in. long, finely hairy beneath. Rootstock: Thick, fleshy. Fruit: A cluster of aromatic, round, pale red speckled berries. Preferred Habitat - Moist woods, thickets, hillsides. Flowering Season - May-July. Distribution - Nova Scotia to Georgia; westward to Arizona and British Columbia.
As if to offer opportunities for comparison to the confused novice, the true Solomon's seal and the so-called false species - quite as honest a plant - usually grow near each other. Grace of line, rather than beauty of blossom, gives them both their chief charm. But the feathery plume of greenish-white blossoms that crowns the false Solomon's seal's somewhat zig-zagged stem is very different from the small, greenish, bell-shaped flowers, usually nodding in pairs along the stem, under the leaves, from the axils of the true Solomon's seal. Later in summer, when hungry birds wander through the woods with increased families, the wild spikenard offers them branching clusters of pale red speckled berries, whereas the latter plant feasts them with blue-black fruit, in the hope that they will drop the seeds miles away.
By clustering its small, slightly fragrant flowers at the end of its stem, the wild spikenard offers a more taking advertisement to its insect friends than its cousin can show. A few flies and beetles visit them; but apparently the less specialized bees, chiefly those of the Ha...